This paper aims to contribute to the development of third-generation activity theorizing. It does so by analysing (inter)acting subjects engaged in joint work at the borders of their respective activity systems. The paper explores these issues theoretically by discussing practices observed in pilot‚Äö-controller interaction in the aviation industry. In this case the way in which work practices are jointly mediated through changes occurring in interacting activity systems are examined. The analysis will show how the changes in elements of the activity systems reveal points of tension and contestation and thus opportunities for development in the everyday interactions between air traffic controllers and airline pilots. In undertaking the analysis, the paper introduces some key questions for consideration when designing interventions in such work environments and contributes to the development of third-generation activity theory.