Stallion harassment and the mating system of horses

Feral horse, Equus caballus, breeding groups, called bands, usually include one but sometimes up to five stallions. We found that mares were loyal to single-stallion (SS) or multistallion (MS) bands or were social dispersers (maverick mares, Mv). The spacing and social behaviour of mares and stallions in single- and multistallion bands was measured. Indices of mare well-being were also measured including activity budgets (feeding: MS>SS = Mv; resting: MSSS), maternal effort in maintaining contact with foals (MS = Mv>SS), parasite levels in faeces (MS>Mv>SS), body condition (MS = Mv